Zeke Yeager

The Ape Titan is an unusual Titan that was first seen within Wall Rose, and potentially a central antagonist in the whole series.

Appearance
The Ape Titan is 17m in stature, larger than the common 15m Titan. While all other Titans seen in the story thus far have been distinctly human in appearance, the Ape Titan possesses more animal-like features. It has apelike features, and most of its body except for the face and torso is covered in fur.

Abilities
The Ape Titan possesses numerous abnormal traits. Most notable are its high intelligence and its ability to speak human language. Possibly as a consequence of this, it is capable of controlling other Titans through a combination of verbal orders and physical intimidation. Additionally, the Ape Titan possesses the strength to throw heavy projectiles great distances with excellent accuracy and the dexterity to scale the Wall Maria.

Story
The Ape Titan first appears when Titans mysteriously appear within Wall Rose even though the Wall remains intact. The Titan displays remarkable intelligence in its encounter with Mike Zakarius. It prevents his escape by killing his horse, speaks the human language, orders other Titans not to eat him so that it can question him about the Three Dimensional Maneuver Gear, and takes the gear for investigation when Mike refuses to respond to it.

The Ape Titan walks past Castle Utgard and starts to climb the Wall Sina.

As the Ape Titan breaks off chunks of the Wall Sina and throws them like missiles at members of the Scouting Legion trapped in Castle Utgard. The first shot kills their horses, while the second kills Rene and Henning.

The Ape Titan is mentioned by Bertholdt and Reiner as an ally to them and that its the mastermind behind the titans attacking the wall.