Zeke Yeager

The Ape Titan is an unusual Titan that was first seen within Wall Rose during it's supposed breach, and is potentially a major antagonist in the overall series.

Appearance
The Ape Titan is 17m in stature, larger than the common 15m Titan. While all other Titans seen in the story thus far have been distinctly human in appearance, the Ape Titan possesses more animal-like features. It has apelike features, and most of its body except for the face and torso is covered in fur.

Abilities
The Ape Titan possesses numerous abnormal traits. Most notable are its high intelligence and its ability to speak human language. Possibly as a consequence of this, it is capable of controlling other Titans through a combination of verbal orders and physical intimidation. Additionally, the Ape Titan possesses the strength to throw heavy projectiles great distances with excellent accuracy and the dexterity to scale the Wall Rose.Another abnormal trait is it's interest in acquiring knowledge about the 3DMG.

Story
The Ape Titan first appears when Titans mysteriously appear within Wall Rose even though the Wall remains intact. The Titan displays remarkable intelligence in its encounter with Mike Zakarius. It prevents his escape by killing his horse, speaks the human language, orders other Titans not to eat him so that it can question him about the Three Dimensional Maneuver Gear, and takes the gear for investigation when Mike refuses to respond to it.

The Ape Titan walks past Castle Utgard and starts to climb the Wall Rose.

As the Ape Titan breaks off chunks of the Wall Rose and throws them like missiles at members of the Scouting Legion trapped in Castle Utgard. The first shot kills their horses, while the second kills Rene and Henning. Afterwards, it summons more Titans to resume the attack on Castle Utgard, appearing to toy with the soldiers trapped inside.

The Ape Titan is mentioned by Ymir as being directly responsible for the appearance of Titans within Wall Rose, with the possibility that it is "testing" humanity's strength. She reveals that locating it was one of Reiner and Bertolt's primary goals, and key to returning to their village. In a later conversation, Bertolt confirms that they consider it important to their mission and hints that they may consider it a potential ally.

Trivia

 * The Titan's appearance resembles the one of the form of a cryptid known as Yeti.
 * It is hinted, but unconfirmed, that it has some method of transforming humans into Titans. The people from Connie Springer's village appear to have been victims of this, and were unique in their ability to function at night.
 * It appears to have limited or no contact with humanity over a long period of time, as it knows nothing about their equipment or whether they even speak the same language.