Source of all living matter

"Long, long ago... when nothing but mere matter existed in this world. As countless masses of some unknown things appeared, disappeared, then appeared again... one of them eventually survived. We now know it as 'life.'"

- Zeke Yeager explains the origin of life

The "source of all living matter" (有機物の起源) is the tentative name attributed to the anomaly which granted Ymir Fritz the Power of the Titans almost two thousand years ago.

Properties
While it is unknown whether or not the "source of all living matter" is indeed the origin of all life on Earth, it is evidently capable of dramatically affecting organic life in its immediate vicinity. A tree which stood over the anomaly in ancient times grew to immense proportions compared to the surrounding flora. Physical contact between Ymir Fritz and the anomaly caused the creation of the Power of the Titans and subsequent connection of living and dead consciousness in the Eldian race through the paths. Zeke Yeager speculates that the nature of the Power of the Titans, the creation of immense and nearly indestructible bodies and a path to a deathless existence, may have resulted from Ymir Fritz's vulnerability and fear of death.

While the anomaly appears sedentary in nature, it has demonstrated some level of physical initiative, actively reaching its limbs towards Ymir as she approached and later materializing out of Eren Yeager's corpse to connect to his severed head on two separate occasions.

History
While fleeing from her Eldian tormentors, Ymir Fritz encountered the life-giving anomaly beneath a colossal tree in the wilderness. Upon physical contact, she was imbued with the Power of the Titans, gaining the immense form of the Founding Titan.

Return to Shiganshina arc
In his journals, Grisha Yeager recounts his conversation with Eren Kruger in which Kruger elaborates on the varying stories surrounding Ymir and the origin of the Titans. While some consider Ymir's power to be of the Devil and others believe her power was a gift from God, Kruger notes that some believe the origin of Ymir's power was the "source of all living matter," though he admits all explanations of such history are ambiguous and ever-changing.

War for Paradis arc
While attempting to come into contact with Zeke Yeager to unlock the power of the Founding Titan, Eren is decapitated by a shot from an anti-Titan rifle wielded by Gabi Braun. Regardless, their contact is achieved and sends the consciousnesses of the two brothers into the paths with Ymir where Eren witnesses the origin of Ymir's power. Upon gaining control of the Founding Titan moments after decapitation, the anomaly manifests from Eren's body and reattaches itself to his head, saving Eren from completely dying. Gabi Braun witnesses this event and later recounts it to her comrades many days later, comparing the anomaly to a "shining centipede". Suspecting the anomaly to be the true source of the Power of the Titans, Gabi anticipates seeing the anomaly once again if Eren were decapitated a second time. Shortly afterwards, as anticipated, the anomaly manifests out of the spine of Eren's Founding Titan after Jean Kirstein decapitates it with explosives, this time on a much larger scale. The colossal anomaly attempts to enter the Founding Titan's severed head but is stopped by Reiner Braun's Armored Titan. Reiner holds the anomaly in place while Armin Arlert transforms into the Colossus Titan, enveloping both Reiner and the anomaly in its explosive transformation.

The anomaly survives the detonation however, so Reiner and Armin moves toward it in an attempt to kill it. However, another explosion reports from the crater behind them, where Eren reemerges in a more conventional -if still colossal- titan body. Armin moves to face him, leaving Reiner to deal with the "shining centipede." Before he could reach it, the serpentine being begins to exude a large amount of gas as it continues writhing, billowing towards the top of Fort Salta. The assembled soldiers quickly determine that this gas is similar to Zeke's spinal fluid cloud, used to turn the inhabitants of Ragako Village into Titans. Shortly after, the remaining "normal" Eldians are transformed into mindless titans, which unusually do not begin to target the remaining humans. Rather, they leap off the mesa, rushing to defend the Source of All Living Matter. As Reiner wrestles with the worm, the titans reach him, tearing into his body. Pieck Finger transforms into the Cart Titan, slowing the creature down as Reiner begins to become overwhelmed. Annie Leonhart tugs it backward using the Female Titan, unimpeded as the horde of titans have not begun to notice her. They swiftly do though, and the three Titan-shifters are assaulted by the horde as they desperately try to slow the "centipede" down.

Trivia

 * The "source of all living matter" bears some resemblance to , an ancient species of aquatic lobopodian worm from the Cambrian period. This resemblance is made more apparent by the appearance of a Hallucigenia sparsa during Zeke Yeager's discussion of the origin of life.
 * While it is not currently clear whether or not the "source of all living matter" is truly sentient, it could be considered the overarching antagonist of Attack on Titan due to its influence on the series' events.

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